Well to me, living in ND the one in the radiator is a fluid warmer and the one in front of the radiator is the cooler. I removed all the stock plumbing that went to them. Replaced all the metal lines with 3/8 trans cooler lines and installed an aftermarket Derale 13011 tranny thermostat off Amazon. Now in order for the torque converter to lock up it needs to be at a certain temp, right. I plumbed the lines so that the transmission uses the heat from the radiator to warm the fluid, then at 180 degrees the Derale thermostat opens and allows fluid to flow through the front cooler. Alot of guys living in colder areas have to put something over the cooler in the winter so the torque converter will lock up and get better gas mileage. With this set up I no longer have to do that and my torque converter locks in -10 degrees. Also increasing flow and allowing for better shifts. You dont have to do this but to me it just made sense.